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The grip stippling and 1 generation of enhancements. Full details are found here : https://thefiringline.com/forums/showthread.php?t=248632

The Millennium Pro pistols offer many improvements over the standard models, but were introduced as enhancements to the originals, not to supplement them. Pro counterparts have now been released for nearly all of the original models. Some features of the Pro line include optional increased magazine capacities, larger and easier to operate controls, a single-action/double-action trigger mechanism on some models with reduced trigger travel and an improved smoother and lighter trigger pull, an enhanced frame, and an improved more ergonometric grip design. Key to the grip upgrade are recesses which Taurus call 'Memory pads', intended to promote a consistent shooting grip from shot to shot. Other Pro improvements include the contrast enhanced Heinie 'Straight-8' sighting system, as well as optional night sights, a loaded chamber indicator which can be felt as well as seen, and a feature labeled 'posi-traction' which consists of an alteration to the number and layout of the slide serrations