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Hi John and welcome. Yes, all 375 Winchester rifles utilize chambers long enough to accept the 2.085 inch 38-55 brass, even though 375 Winchester brass is only 2.020 inches long. Cheers and stay safe!

John, the information below is according to Buffalo bore........ Our 38-55 ammo is also designed to work in ALL 375 Winchester chambered rifles. ALL means ALL. Winchester, Savage, and Marlin all chambered rifles in 375 Winchester. All 375 Winchester rifles utilize chambers long enough to accept the 2.085 inch 38-55 brass, even though 375 Winchester brass is only 2.020 inches long.......... Im not finding any information regarding this specific ammo. So I'd recommend contacting Winchester customer service for a definite answer. If interested here's a link you can use to message them. Sorry can't give a definite answer but hope this helps some, link here.................................... http://www.winchester.com/Customer-Support/Pages/Contact-Us.aspx